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[13] In 1955, Heinz Fraenkel-Conrat and Robley Williams showed that purified TMV RNA and its capsid (coat) protein assemble by themselves to functional viruses, indicating that this is the most stable structure (the one with the lowest free energy)

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The development and significance of Holmes findings, within the context of agriculture and plant pathology in the United States in the early 20th century, is a key area of interest in revealing the history of TMV and plant pathology in the United States