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All statistical analyses were performed using R Statistical Software (v4.1.3) ( Aligned reads were then compiled into a single bedfile for each experimental group using BEDOPS toolkit (2.4.41) 42

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First, the periplasmic fractions were bound to HisPur Ni-NTA Resin (Thermo 88222) and then washed sequentially with TBS (20 mM Tris, 150 mM NaCl) containing 20 mM and 40 mM imidazole, followed by elution with TBS containing 500 mM imidazole