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Growth of LAB under acid, bile salts, and NaCl The LAB strains were inoculated (2%, v/v) into MRS broth and the cells were harvested by centrifugation after cultured for 18 h

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The frequently studied ligand environments around the Mn center were salen derivatives, nitrogen-centered ligands, cyclic polyamines, carboxylate/ amino-carboxylate ligands, porphyrins, peptides, phthalocyanines and other ligands (Forman and Zhang 2021)