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Phenylpropanoid metabolism is one of the most important secondary metabolic pathways in plants

This process begins with the oxidation of 5-methylcytosine to 5-hydroxymethylcytosine, followed by further oxidation to 5-formylcytosine and 5-carboxylcytosine, with the oxidized intermediates eventually removed through the base excision repair mechanism, restoring the DNA to an unmethylated state

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This results in impaired mitochondrial integrity, impaired defense against ROS and imbalances in protein homeostasis

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