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Chen YL, Shi L, Agbo F, Yong SH, Tan PS, Ngounou Wetie AG

(2000), in Life Sciences , extended this work into matrix metalloproteinase (MMP) biology, finding that GHK-Cu exposure elevated MMP-2 expression in fibroblast cultures while simultaneously upregulating tissue inhibitors of metalloproteinases (TIMP-1 and TIMP-2).[6] The significance here is the pairing: GHK-Cu appears to promote a coordinated remodeling response controlled matrix breakdown balanced by inhibitory regulation rather than simply driving one-directional collagen accumulation

It works primarily through actin regulation, which gives it a broad reach across different tissue types

Tight Junctions as Modulators of Intestinal Integrity and Paracellular Translocation The paracellular transport of important molecules across the intestinal epithelium is regulated by transmembrane and intracellular proteins that interlink to form TJs