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Moreover, Gln is an essential substrate for the immune system, and prolonged use of Gln metabolic inhibitors might weaken the anti-tumor immune response ( 2 The role of Gln metabolism in tumor Emerging evidence shows that Gln metabolism is involved in tumor progression in two specific ways ( Figure 1 )

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To shed more light on the identity of the substrate of arGSTs, we performed labelling experiments with H 2 18 O, exploiting the same principle of 18 O incorporation as described earlier

Others include ICOS, CD27, CD40, and OX40

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