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In contrast, although the proportion of H2AX-positive cells increased for both HCT116 GSTA4 (1.67 2.42%) and HCT116 cells (3.69 6.00%) after treatment with oxaliplatin for 48 h compared with untreated controls, no significant difference in the proportion of H2AX-positive cells was seen between HCT116 GSTA4 and HCT116 cells ( Supplementary Figures 3A, B , P = 0.618), indicating that the inactivation of GSTA4 has limited effect on low-dose oxaliplatin-induced crosslinking of DNA

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Therefore, the depletion of gliN or the overexpression of gtmA can alleviate the production of gliotoxin and promote the detoxification of gliotoxin, respectively