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doi:10.4021/JEM.V3I1-2.157 Heffernan M, Summers RJ, Thorburn A, et al

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While Nirenberg and team were using binding assays to try to capture the RNA-amino acid associations, Khorana approached it differently by synthesizing repeating dinucleotide and trinucleotide RNA polymers and seeing which amino acids ended up linked together in the resulting protein chain