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For GHK-Cu specifically, the logic is sound even without direct evidence: cumulative copper exposure is lower with reduced frequency, and the gene expression changes initiated during loading persist during maintenance

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He then went on to estimate their population, despite expressing misgivings about the accuracy of the estimate, at from 200,000 to 300,000 people.[26] From 1893 and Hodges initial estimate, archaeologists have been generally driving the numbers down