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Similarly, in an inducible rat bead model of glaucoma, metabolic profiles were demonstrated to be dysregulated in the whole retina, optic nerve and superior colliculus, occurring at a timepoint when IOP was high but before detectable retinal ganglion cell degeneration ( Metabolomics analyses of whole glaucomatous retina in animal models have identified potential changes to several metabolic pathways including alanine, aspartate and glutamate metabolism, D-glutamine and D-glutamate metabolism, and arginine biosynthesis (36)