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Most of the PAPs have PHR1 binding sites in their promoters and are responsive to Pi deficiency ( osarf12 , elevated the expression of 4 PAPs genes OsPAP9b, OsPAP10a, OsPAP10c , and OsPAP27a enhanced APase activity, suggesting that ARF12 influenced transcript abundance from PAP genes in P sufficient and deficient conditions (Wang et al., 2014b)